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193 These subunits can directly interact with the activated GPCR or do so via intermediary proteins such as insulin receptor substrate

One proposed mechanism involves the regulation of chromatin-bound copper ions, leading to oxidative DNA damage and fragmentation that triggers apoptotic pathways (289, 290)

Their dual role is exemplified by their ability to inactivate mTORC1 at high concentrations, triggering protective processes such as mitophagy, whereas at lower concentrations, they can activate mTORC1 and promote cell survival pathways