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All authors have read and approved the final manuscript

The four polypeptide chains are bound to each other by salt bridges, forming a tetrameric quaternary structure

Next, we examined the enzymatic activities of AsGGT1, AsGGT2, and AsGGT3 toward -glutamyl- S -allyl-L-cysteine and -glutamyl- S -allyl-L-cysteine sulfoxide, which are two possible biosynthetic intermediates in alliin biosynthesis, as potential -glutamyl donor substrates ( Figures 3 and 4 )