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Limitations of the study The study enrolled patients with active RA, with a significant proportion receiving dual or triple DMARDs therapy

In 2001, two research groups published papers reporting that, under normoxia, the HIF-1 subunit is targeted by PHDs and is rapidly destroyed via a mechanism that involves ubiquitylation by the von Hippel-Lindau tumor suppressor (pVHL) E3 ligase complex [43,44]